The Natural Communities of Virginia
Classification of Ecological Community Groups
SECOND APPROXIMATION (Version 2.2)
Mesic Mixed Hardwood Forests
These mixed hardwood forests are widespread in mesic to submesic, infertile habitats throughout the Coastal Plain and Piedmont, and rarely at low elevations in the mountains. Forests in this group occupy mesic uplands, ravines, lower slopes, and well-drained “flatwoods” on acidic, relatively nutrient-poor soils. The most typical overstories contain mixtures of American beech (
Fagus grandifolia), oaks (
Quercus spp., varying by region), tulip-poplar (
Liriodendron tulipifera), and hickories (
Carya spp.), but a wide variety of hardwood associates occur. American hornbeam (
Carpinus caroliniana ssp.
caroliniana and ssp.
virginiana), flowering dogwood (
Cornus florida), American strawberry-bush (
Euonymus americanus) and, in eastern Virginia, American holly (
Ilex opaca var. opaca) are prominent understory plants. In mesic “flatwoods” of the southeastern Virginia Coastal Plain, silky camellia (
Stewartia malacodendron) and big-leaf snowbell (
Styrax grandifolius) are characteristic small trees. These communities lack the lush herbaceous layers of Basic Mesic Forests, although species such as Christmas fern (
Polystichum acrostichoides), New York fern
(
Thelypteris noveboracensis), and white wood aster (
Eurybia divaricata, =
Aster divaricatus) may form moderately dense populations. Along with Chistmas fern, downy rattlesnake-plantain (
Goodyera pubescens), Virginia heartleaf (
Hexastylis virginica), and partridge-berry (
Mitchella repens) are frequent evergreen herbs in mesic mixed hardwood forests. The name “Southern Mixed Hardwood Forest” has been applied to some Coastal Plain representatives of this group. Although mesic mixed hardwood forests still cover sizeable areas east of the mountains in Virginia, their extent and compositional integrity have been reduced by repeated logging. Several distinct community types are represented in this widespread group.
References: Abrams and Copenheaver (1999), Adams et al. (2003), Coulling (1999), Crouch (1990),
DeWitt and Ware (1979), Fleming (2002a), Fleming (2002b), Frost and Musselman (1987), McCoy and Fleming (2000), Monette and Ware (1983),
Plunkett and Hall (1995), Walton et al. (2001), Ware (1970), Ware (1978), Ware (1991), Wolff and Ware (1994).
Click on the images below to open a larger image in a separate window.
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The evergreen Christmas Fern (Polystichum acrostichoides, in foreground) is one
of the most characteristic and abundant herbs of mesic mixed hardwood forests. Riverbend Park, Fairfax County.
Photo: Gary P. Fleming
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American beech (Fagus grandifolia), American holly (Ilex opaca var. opaca ), and white oak
(Quercus alba) in a southern Piedmont Mesic Mixed Hardwood Forest. Pocahontas State Park, Chesterfield County.
Photo: Gary P. Fleming.
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REPRESENTATIVE COMMUNITY TYPES:
This group is represented by more than 100 quantitative plot samples, most of them supporting
the first two classified types. Representatives of this group in the Embayed Region, although clearly distinct, need more
documentation. The Piedmont / Northern Coastal Plain Type encompasses a considerable gradient in apparent site moisture potential,
from that of very mesic ravine bottoms to that of submesic, convex upper slopes and crests. Nevertheless, attempts to split this
unit into two or more types or subtypes have proven difficult to justify on the basis of floristic composition, despite the
availability of ample quantitative data. Click on any highlighted CEGL code below to view the global USNVC description provided by NatureServe Explorer.
- Fagus grandifolia – Liriodendron tulipifera – Quercus ( alba , rubra ) / Polystichum acrostichoides Forest
Mesic Mixed Hardwood Forest (Piedmont / Northern Coastal Plain Type)
USNVC: = CEGL006075
Global/State Ranks:
GNR/S5
- Fagus grandifolia – Quercus alba / Ilex opaca – ( Oxydendrum arboreum ) / Vitis rotundifolia Forest
Mesic Mixed Hardwood Forest (Coastal Plain / Southern Piedmont Type)
USNVC:
~ CEGL008465
Global/State Ranks:
G3G4/SNR
- Fagus grandifolia – Quercus ( alba , michauxii , pagoda ) / Stewartia malacodendron Forest (PROVISIONAL)
Mesic Mixed Hardwood Forest (Southeastern Embayed Region Type)
USNVC:
= CEGL007211
State/Global Ranks: G3/S2
