Montane Mixed Oak and Oak - Hickory Forests
This group contains relatively diverse, mixed oak and oak-hickory forests of submesic to subxeric mountain slopes and crests occurring mostly between about 600 m (2,000 ft) and 1,200 m (4,000 ft) elevation.
Montane mixed oak forests cover extensive areas in western Virginia and generally occupy intermediate positions along major environmental gradients such as soil moisture, soil fertility, and elevation. Soils occupied by montane mixed oak forests are typically more acidic than those of the true montane oak-hickory forests. Overstory composition contains mixtures of chestnut oak (Quercus montana), northern red oak (Quercus rubra), and white oak (Quercus alba). Overstory associates vary with geography and site conditions, but often include sweet birch (Betula lenta), magnolias (Magnolia acuminata and Magnolia fraseri), sourwood (Oxydendrum arboreum), hickories (Carya spp.), red maple (Acer rubrum), tulip-poplar (Liriodendron tulipifera), and white pine (Pinus strobus). The understories of mixed oak communities usually contain a substantial component of heaths, but also contain many non-ericaceous species such as witch-hazel (Hamamelis virginiana), striped maple (Acer pensylvanicum), maple-leaved viburnum (Viburnum acerifolium), mountain holly (Ilex montana), buffalo-nut (Pyrularia pubera), and hazelnuts (Corylus cornuta var. cornuta and Corylus americana). The herbaceous component is relatively diverse, but often patchy and composed of both acidophiles and species characteristic of moderately fertile soils, including New York fern (Thelypteris noveboracensis), galax (Galax urceolata), Curtis' goldenrod (Solidago curtisii), white wood aster (Eurybia divaricata), indian cucumber-root (Medeola virginiana), squawroot (Conopholis americana), halberd-leaved yellow violet (Viola hastata), white clintonia (Clintonia umbellulata), devil's-bit (Chamaelirium luteum), mountain golden alexanders (Zizia trifoliata), and American lily-of-the-valley (Convallaria majuscula).
Montane oak-hickory forests reach maximal importance on base-rich igneous, metamorphic, and subcalcareous sedimentary rocks. Two types occur throughout western Virginia: a submesic to mesic, rich type with a notably lush herb layer, and a drier, more acidic type with a diverse, often graminoid-dominated herb layer. Northern red oak, white oak, red hickory (Carya ovalis), and shagbark hickory (Carya ovata) are typical co-dominant trees, although in most stands oaks attain greater importance in the overstory than do hickories (Carya spp.), which often reach maximal density and cover in subcanopy strata. The chestnut oak is important only in the drier type, while mesophytic trees such as basswood (Tilia americana var. americana and var. heterophylla), bitternut hickory (Carya cordiformis), and sugar maple (Acer saccharum var. saccharum) are frequent associates in the rich type. The shrub layer of these communities is often sparse, and herbaceous composition varies with geography and site conditions. A widespread compositional variant on moist sites features extensive, nearly monospecific colonies of interrupted fern (Osmunda claytoniana). Fertile sites often support a diverse herbaceous flora, including such nutrient-demanding forbs as purple giant hyssop (Agastache scrophulariifolia), basil bee-balm (Monarda clinopodia), green-head coneflower (Rudbeckia laciniata var. laciniata), pale-leaf woodland sunflower (Helianthus strumosus), richweed (Collinsonia canadensis), yellow jewelweed (Impatiens pallida), black bugbane (Cimicifuga racemosa), starry campion (Silene stellata), stout goldenrod (Solidago squarrosa), hairy-jointed meadow-parsnip (Thaspium barbinode), and Appalachian meadowrue (Thalictrum coriaceum). The luxuriance of such herb layers rivals that of the Rich Cove and Slope Forests. The drier type of montane oak-hickory forest features patch-dominance by Pennsylvania sedge (Carex pensylvanica), Porter's reedgrass (Calamagrostis porteri ssp. porteri), wavy hairgrass (Deschampsia flexuosa var. flexuosa), and xerophytic forbs.
Communities of this group are transitional to Northern Red Oak Forests at higher elevations and on poorer sites and to several other oak and oak-hickory forests at their lower-elevation limits. They are distinguished from both Basic Oak-Hickory Forests and Acidic Oak-Hickory Forests by their restriction to elevations generally > 600 m (2,000 ft) and a floristic composition that lacks many prominent low-elevation species (e.g., eastern redbud [Cercis canadensis var. canadensis] and flowering dogwood [Cornus florida]). They are distinguished from Oak / Heath Forests by their far greater diversity of understory and herbaceous plants. Many Montane Mixed Oak and Oak-Hickory stands represent vegetation formerly dominated or co-dominated by American chestnut (Castanea dentata), before mature individuals of this species were decimated by the introduced chestnut blight (Cryphonectria parasitica) in the early 20th century. Hickories (Carya spp.) are thought to have benefited greatly from the removal of American chestnut from the overstory, and their persistence and continued recruitment in contemporary oak-hickory forests may reflect fire exclusion in recent decades.References: Adams and Stephenson (1983), Coulling and Rawinski (1999), Fleming (2007), Fleming and Coulling (2001), Fleming and Moorhead (2000), Johnson and Ware (1982), McCormick and Platt (1980), Rawinski et al . (1994), Rawinski et al . (1996), Stephenson (1982a), Stephenson (1982b), Stephenson and Adams (1991).
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Southern Appalachian Montane Mixed Oak forest co-dominated by northern red oak (Quercus rubra) and chestnut oak (Quercus montana), with understory of sourwood (Oxydendrum arboreum) and striped maple (Acer pensylvanicum). North slope of Hurricane Mountain, Smyth Co. (George Washington and Jefferson National Forests). Photo: Gary P. Fleming
|The lush herb layers of some montane oak-hickory forests rival those of the rich cove and slope forests. Summit of the northern Blue Ridge near the headwaters of Entry Run, Greene County (Shenandoah National Park). Photo: Gary P. Fleming|
|Emerging fronds of interrupted fern (Osmunda claytoniana) in a montane oak-hickory forest on Thunder Hill, Bedford County (George Washington and Jefferson National Forests). Photo: Gary P. Fleming|
|Montane oak-hickory forest on the crest of the northern Blue Ridge near Humpback Mountain, Nelson County (Blue Ridge Parkway). Black bugbane (Cimicifuga racemosa, flowering) dominates the dense herb layer. Gary P. Fleming / © DCR Natural Heritage.|
|White oak (Quercus alba), red hickory (Carya ovalis), and Appalachian meadowrue (Thalictrum coriaceum) dominate a stand of montane oak-hickory forest on The Nature Conservancy's Warm Springs Mountain preserve, Bath County. Gary P. Fleming / © DCR Natural Heritage.|
This group is represented by more than 240 plot samples from most counties in western Virginia and a few outlying Piedmont sites (map). The classification of six community types is well supported by several large regional analyses of plot data. However, the Southern Appalachian representatives of the group need additional inventory to determine their full within-state distribution. Click on any highlighted CEGL code below to view the global USNVC description provided by NatureServe Explorer.
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